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A non-zero number of seeds germinated after 2 years instead of just 1 in Bruna 2002 (Fig 4). E.g., it looks like in continuous forest, and additional ~4% of seeds germinated in the second year. One way to model this could be to add another discrete class, seed that has age related transitions into sdlg.
Bruna, Emilio M. “Effects of Forest Fragmentation on Heliconia Acuminata Seedling Recruitment in Central Amazonia.” Oecologia 132, no. 2 (2002): 235–43. https://doi.org/10.1007/s00442-002-0956-y.
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